Moots,
Last night I put a nuc in my home made ulster OH to do a school bee presentation. It has a open screen bottom and an 3/4" opening with an adjustable open/screen/QE devise. I put the OH on my bar top to have it ready for the presentation. Late last night the bees were running and started removing open brood from their cells. My wife pointed out that the entrance was packed with bees behind the screen. I realized that putting it on the bar directly caused it to be sealed air tight. I raised it up on the 4 corners and after a while they calmed.
I recommend you check out how much ventilation on your Oh. I am pretty sure this is what happened to my large OH. The bees had closed up the top vents during the winter and I noticed when I cleaned it out that they did open one of them. Maybe too late. I think the dead bees blocked the feeder entrance and the main entrance, blocking most of the air flow.
Let me know if this could bee your problem also.
Jim