I took this frame out I then replaced it with a complete new frame & foundation.
It really seems to me that the right end of frame (1/4) have got condensation moisture and mould have developed in new part of wax and in walls of cells.
That is the reason why I put a foundation against the wall for winter. If the edge frame is old mould attaches very easily to those combs. In nature same happens and bees tear rotten spoiled wax away. Mold eat the layers of cocoon silk in the cell. That is why old cell walls are like swollen. Mould may be white or blue in our country. If dead bees are so plenty that they reach the lowest comb, those parts will be ruined by blue mould. BleebShit happens, we say.
You renewed it. It is well done. I have had those more than enough. I kept combs too long. When you sniff the full of these frames odor is not fresh like in new combs. This odor goes to the taste of honey.