In my vast season and a half of keeping....lol.... I have noticed something.
Give the bees foundationless frames in the brood area and they build consistent, straight, 'perfect' comb. Give this same hive empty frame in a super and the comb is more often wonky. Different thickness of comb, every spare cm of space is used for storage or the management of the honey. Sometimes I even wonder if they design it such that it promotes unique air flow within the hives.
My observation was consistent with Carnolians, Italians, and my local mutts.